Codexis, Inc.CDXSNASDAQ
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| Holder | Shares | Date | Change | % Held |
|---|---|---|---|---|
| Physician Wealth Advisors, Inc. | -- | Dec 31, 25 | -442 | -- |
| NISA INVESTMENT ADVISORS, LLC | -- | Dec 31, 25 | -9.14K | -- |
| CRUX WEALTH ADVISORS | -- | Dec 31, 25 | -13.41K | -- |
| COMERICA BANK | -- | Dec 31, 25 | -972 | -- |
| Opaleye Management Inc. | 12.79M | Sep 30, 25 | +195.00K | -- |
| BlackRock, Inc. | 7.14M | Sep 30, 25 | +8.01K | -- |
| MILLENNIUM MANAGEMENT LLC | 4.07M | Sep 30, 25 | +3.95M | -- |
| Nantahala Capital Management, LLC | 4.03M | Sep 30, 25 | -- | -- |
| Telemark Asset Management, LLC | 4.00M | Sep 30, 25 | -- | -- |
| AMERIPRISE FINANCIAL INC | 2.77M | Sep 30, 25 | +222.84K | -- |
| GEODE CAPITAL MANAGEMENT, LLC | 1.99M | Sep 30, 25 | +14.54K | -- |
| Pier Capital, LLC | 1.89M | Sep 30, 25 | -23.93K | -- |
| STATE STREET CORP | 1.54M | Sep 30, 25 | -72.58K | -- |
| Empire Financial Management Company, LLC | 1.46M | Sep 30, 25 | +1.17M | -- |
| JACOBS LEVY EQUITY MANAGEMENT, INC | 1.16M | Sep 30, 25 | -13.29K | -- |
| Nuveen, LLC | 920.12K | Sep 30, 25 | +619.27K | -- |
| FRANKLIN RESOURCES INC | 756.16K | Sep 30, 25 | +308.63K | -- |
| GOLDMAN SACHS GROUP INC | 724.25K | Sep 30, 25 | +275.49K | -- |
| NORTHERN TRUST CORP | 645.27K | Sep 30, 25 | -13.88K | -- |
| PRESCOTT GROUP CAPITAL MANAGEMENT, L.L.C. | 635.54K | Sep 30, 25 | -65.05K | -- |
| DIMENSIONAL FUND ADVISORS LP | 603.62K | Sep 30, 25 | +31.48K | -- |
| FMR LLC | 584.37K | Sep 30, 25 | -4.03M | -- |
| Private Advisor Group, LLC | 561.41K | Sep 30, 25 | +38.50K | -- |
| TUDOR INVESTMENT CORP ET AL | 537.83K | Sep 30, 25 | +537.83K | -- |
| BALYASNY ASSET MANAGEMENT L.P. | 496.79K | Sep 30, 25 | +496.79K | -- |
| ROYAL BANK OF CANADA | 410.13K | Sep 30, 25 | -4.29K | -- |
| RENAISSANCE TECHNOLOGIES LLC | 310.49K | Sep 30, 25 | -445.00K | -- |
| PERKINS CAPITAL MANAGEMENT INC | 307.70K | Sep 30, 25 | -4.30K | -- |
| STIFEL FINANCIAL CORP | 278.34K | Sep 30, 25 | +238.35K | -- |
| Squarepoint Ops LLC | 276.36K | Sep 30, 25 | +259.26K | -- |
| CITADEL ADVISORS LLC | 269.93K | Sep 30, 25 | +158.08K | -- |
| PLATINUM INVESTMENT MANAGEMENT LTD | 264.46K | Sep 30, 25 | -51.92K | -- |
| CHARLES SCHWAB INVESTMENT MANAGEMENT INC | 250.73K | Sep 30, 25 | +6 | -- |
| FEDERATED HERMES, INC. | 234.25K | Sep 30, 25 | -638.57K | -- |
| BARCLAYS PLC | 224.49K | Sep 30, 25 | -29.15K | -- |
| Mink Brook Asset Management LLC | 220.00K | Sep 30, 25 | -101.71K | -- |
| Bank of New York Mellon Corp | 208.25K | Sep 30, 25 | +449 | -- |
| MORGAN STANLEY | 196.63K | Sep 30, 25 | -283.23K | -- |
| D. E. Shaw & Co., Inc. | 173.81K | Sep 30, 25 | +173.81K | -- |
| Cubist Systematic Strategies, LLC | 147.38K | Sep 30, 25 | +147.38K | -- |
| WHITE PINE CAPITAL LLC | 138.19K | Sep 30, 25 | +438 | -- |
| Man Group plc | 133.84K | Sep 30, 25 | +133.84K | -- |
| Diametric Capital, LP | 129.87K | Sep 30, 25 | +129.87K | -- |
| Invesco Ltd. | 123.14K | Sep 30, 25 | +27.06K | -- |
| ALLIANCEBERNSTEIN L.P. | 118.13K | Sep 30, 25 | -- | -- |
| RHUMBLINE ADVISERS | 111.82K | Sep 30, 25 | -23.74K | -- |
| BRIDGEWAY CAPITAL MANAGEMENT, LLC | 100.91K | Sep 30, 25 | -- | -- |
| STRS OHIO | 99.10K | Sep 30, 25 | -16.20K | -- |
| Stonepine Capital Management, LLC | 88.26K | Sep 30, 25 | +88.26K | -- |
| LPL Financial LLC | 87.67K | Sep 30, 25 | -93.00K | -- |
| MARSHALL WACE, LLP | 87.06K | Sep 30, 25 | -15.31K | -- |
| AQR CAPITAL MANAGEMENT LLC | 84.07K | Sep 30, 25 | -487 | -- |
| WELLS FARGO & COMPANY/MN | 76.58K | Sep 30, 25 | +32.39K | -- |
| ENVESTNET ASSET MANAGEMENT INC | 67.72K | Sep 30, 25 | -685 | -- |
| ACADIAN ASSET MANAGEMENT LLC | 60.37K | Sep 30, 25 | -21.67K | -- |
| CITIGROUP INC | 56.04K | Sep 30, 25 | +24.77K | -- |
| OXFORD ASSET MANAGEMENT LLP | 54.04K | Sep 30, 25 | +54.04K | -- |
| JANE STREET GROUP, LLC | 50.26K | Sep 30, 25 | +50.26K | -- |
| PARADIGM CAPITAL MANAGEMENT INC/NY | 50.00K | Sep 30, 25 | -- | -- |
| BANK OF AMERICA CORP /DE/ | 49.75K | Sep 30, 25 | -3.59K | -- |
| Brevan Howard Capital Management LP | 48.73K | Sep 30, 25 | +6.71K | -- |
| Russell Investments Group, Ltd. | 48.44K | Sep 30, 25 | +10.41K | -- |
| PUBLIC EMPLOYEES RETIREMENT SYSTEM OF OHIO | 46.71K | Sep 30, 25 | -- | -- |
| PRICE T ROWE ASSOCIATES INC /MD/ | 44.27K | Sep 30, 25 | +3.77K | -- |
| COMMONWEALTH EQUITY SERVICES, LLC | 41.69K | Sep 30, 25 | -1.12K | -- |
| INTECH INVESTMENT MANAGEMENT LLC | 41.03K | Sep 30, 25 | -7.36K | -- |
| MIRAE ASSET GLOBAL ETFS HOLDINGS Ltd. | 38.74K | Sep 30, 25 | -6.86K | -- |
| JPMORGAN CHASE & CO | 38.34K | Sep 30, 25 | -3.77K | -- |
| ADVISOR GROUP HOLDINGS, INC. | 37.06K | Sep 30, 25 | +16.43K | -- |
| PRELUDE CAPITAL MANAGEMENT, LLC | 35.93K | Sep 30, 25 | +35.93K | -- |
| BLAIR WILLIAM & CO/IL | 35.29K | Sep 30, 25 | -1.54K | -- |
| MetLife Investment Management, LLC | 35.28K | Sep 30, 25 | -15.07K | -- |
| MANUFACTURERS LIFE INSURANCE COMPANY, THE | 29.47K | Sep 30, 25 | -309 | -- |
| TWO SIGMA ADVISERS, LP | 27.50K | Sep 30, 25 | -18.50K | -- |
| Quadrature Capital Ltd | 26.93K | Sep 30, 25 | +26.93K | -- |
| Dynamic Technology Lab Private Ltd | 24.11K | Sep 30, 25 | +24.11K | -- |
| TWO SIGMA INVESTMENTS, LP | 22.75K | Sep 30, 25 | -10.90K | -- |
| Police & Firemen's Retirement System of New Jersey | 19.44K | Sep 30, 25 | -- | -- |
| HRT FINANCIAL LP | 18.81K | Sep 30, 25 | +18.81K | -- |
| SUSQUEHANNA INTERNATIONAL GROUP, LLP | 18.78K | Sep 30, 25 | -83.45K | -- |
| XTX Topco Ltd | 17.30K | Sep 30, 25 | -30.19K | -- |
| ProShare Advisors LLC | 17.21K | Sep 30, 25 | -2.67K | -- |
| NEW YORK STATE COMMON RETIREMENT FUND | 16.89K | Sep 30, 25 | -- | -- |
| BNP PARIBAS ARBITRAGE, SNC | 16.61K | Sep 30, 25 | +8.04K | -- |
| AIGEN INVESTMENT MANAGEMENT, LP | 15.83K | Sep 30, 25 | +15.83K | -- |
| VOYA INVESTMENT MANAGEMENT LLC | 14.84K | Sep 30, 25 | -8.28K | -- |
| PUBLIC EMPLOYEES RETIREMENT ASSOCIATION OF COLORADO | 13.64K | Sep 30, 25 | -- | -- |
| FIDUCIARY TRUST CO | 13.62K | Sep 30, 25 | -- | -- |
| State of Wyoming | 13.10K | Sep 30, 25 | +2.66K | -- |
| Jump Financial, LLC | 10.61K | Sep 30, 25 | +10.61K | -- |
| SEI INVESTMENTS CO | 10.54K | Sep 30, 25 | -- | -- |
| VICTORY CAPITAL MANAGEMENT INC | 10.07K | Sep 30, 25 | +10.07K | -- |
| Tower Research Capital LLC (TRC) | 9.97K | Sep 30, 25 | +24 | -- |
| Legal & General Group Plc | 8.50K | Sep 30, 25 | -- | -- |
| Ameritas Investment Partners, Inc. | 8.43K | Sep 30, 25 | -- | -- |
| CALIFORNIA STATE TEACHERS RETIREMENT SYSTEM | 5.93K | Sep 30, 25 | +3 | -- |
| SRS Capital Advisors, Inc. | 3.53K | Sep 30, 25 | +3.53K | -- |
| AMALGAMATED BANK | 2.78K | Sep 30, 25 | -- | -- |
| CIBC Private Wealth Group, LLC | 1.80K | Sep 30, 25 | -- | -- |
| DEUTSCHE BANK AG\ | 1.65K | Sep 30, 25 | -27.07K | -- |
| Sterling Capital Management LLC | 1.20K | Sep 30, 25 | +631 | -- |
| Optiver Holding B.V. | 1.15K | Sep 30, 25 | +1.15K | -- |
| Rockefeller Capital Management L.P. | 1.00K | Sep 30, 25 | +1.00K | -- |
| MARK SHEPTOFF FINANCIAL PLANNING, LLC | 500 | Sep 30, 25 | -- | -- |
| Global Retirement Partners, LLC | 350 | Sep 30, 25 | -- | -- |
| EverSource Wealth Advisors, LLC | 342 | Sep 30, 25 | -- | -- |
| SBI Securities Co., Ltd. | 322 | Sep 30, 25 | -57 | -- |
| Daiwa Securities Group Inc. | 269 | Sep 30, 25 | -- | -- |
| ROTHSCHILD INVESTMENT LLC | 205 | Sep 30, 25 | -- | -- |
| MAI Capital Management | 138 | Sep 30, 25 | -- | -- |
| PNC FINANCIAL SERVICES GROUP, INC. | 77 | Sep 30, 25 | -879 | -- |
| Allworth Financial LP | 3 | Sep 30, 25 | +3 | -- |
| HUNTINGTON NATIONAL BANK | 2 | Sep 30, 25 | -- | -- |
| NORTHWESTERN MUTUAL WEALTH MANAGEMENT CO | 1 | Sep 30, 25 | -- | -- |
| SouthState Corp | -- | Sep 30, 25 | -200 | -- |
| Parallel Advisors, LLC | -- | Sep 30, 25 | -200 | -- |
| Casdin Capital, LLC | -- | Sep 30, 25 | -7.20M | -- |
| RAYMOND JAMES FINANCIAL INC | -- | Sep 30, 25 | -400 | -- |
| CLOUGH CAPITAL PARTNERS L P | -- | Sep 30, 25 | -69.00K | -- |
| Inspire Investing, LLC | -- | Sep 30, 25 | -162.91K | -- |
| Skopos Labs, Inc. | -- | Sep 30, 25 | -12 | -- |
| GROUP ONE TRADING, L.P. | -- | Sep 30, 25 | -100 | -- |
| CIBC WORLD MARKETS CORP | 10.09K | Jun 30, 25 | -7.79K | -- |
| Nuveen Asset Management, LLC | 1.81M | Dec 31, 24 | +283.76K | -- |
| Tallon Kerry Patrick | 403.67K | Dec 31, 24 | +403.67K | -- |
| nVerses Capital, LLC | 1.00K | Sep 30, 24 | +1.00K | -- |
| BlackRock Inc. | 6.79M | Jun 30, 24 | +174.70K | -- |
| Kent Lake Capital LLC | 1.30M | Jun 30, 24 | +670.60K | -- |
| Point72 Middle East FZE | -- | Dec 31, 23 | -48.92K | -- |
| Trellus Management Company, LLC | 220.00K | Sep 30, 23 | -31.46K | -- |
| JACOB ASSET MANAGEMENT OF NEW YORK LLC | 131.02K | Sep 30, 23 | -67.45K | -- |
| Tekla Capital Management LLC | 69.13K | Jun 30, 23 | -- | -- |
| BerganKDV Wealth Management, LLC | 3.50K | Jun 30, 23 | -- | -- |
| EATON VANCE MANAGEMENT | 150.34K | Sep 30, 22 | -- | -- |
| AlphaCrest Capital Management LLC | 70.97K | Sep 30, 22 | +70.97K | -- |
| Jefferies Group LLC | 14.51K | Sep 30, 22 | +858 | -- |
| Parian Global Management LP | 8.80K | Sep 30, 22 | +8.80K | -- |
| Private Capital Group, LLC | -- | Jun 30, 22 | -368 | -- |
| PARAMETRIC PORTFOLIO ASSOCIATES LLC | 28.77K | Mar 31, 22 | -12.53K | -- |
| Amalgamated Financial Corp. | 9.82K | Mar 31, 22 | +9.82K | -- |
| KNOTT DAVID M | 10.00K | Sep 30, 21 | +10.00K | -- |
| COASTAL INVESTMENT MANAGEMENT, L.P. | -- | Sep 30, 21 | -39.00K | -- |
| Pratt Collard Advisory Partners LLC | 68.05K | Jun 30, 21 | +300 | -- |
| Aperio Group, LLC | 15.02K | Dec 31, 20 | +1.45K | -- |
| VIRTUS ALTERNATIVE INVESTMENT ADVISERS, INC. | 470 | Dec 31, 20 | -453 | -- |
| Cornerstone Advisors, Inc. | -- | Sep 30, 20 | -71 | -- |
| Leap Investments LP | -- | Jun 30, 20 | -13.60K | -- |
| LADENBURG THALMANN FINANCIAL SERVICES INC. | 1.71K | Dec 31, 19 | -21 | -- |
| Advisor Group, Inc. | 1.13K | Dec 31, 19 | -1.27K | -- |
| BOSTON ADVISORS LLC | 29.36K | Sep 30, 19 | +8.74K | -- |
| Cortina Asset Management, LLC | -- | Jun 30, 19 | -183.83K | -- |
| TEACHERS ADVISORS, LLC | 98.64K | Mar 31, 19 | +4.67K | -- |
| TIAA CREF INVESTMENT MANAGEMENT LLC | 85.32K | Mar 31, 19 | -2.85K | -- |
| Virtu KCG Holdings LLC | -- | Jun 30, 17 | -15.58K | -- |
| BlackRock Institutional Trust Company, N.A. | 1.24M | Dec 31, 16 | +14.18K | -- |
| BlackRock Fund Advisors | 1.11M | Dec 31, 16 | +192.63K | -- |
| BlackRock Investment Management, LLC | 123.64K | Dec 31, 16 | -- | -- |
| BLACKROCK ADVISORS LLC | 12.25K | Dec 31, 16 | -86 | -- |
| UBS AG | 4.71K | Sep 30, 14 | +8 | -- |
| CONNOR CLARK & LUNN INVESTMENT MANAGEMENT LTD | 38.40K | Dec 31, 13 | -800 | -- |
| KNIGHT CAPITAL GROUP LLC | 26.56K | Jun 30, 13 | +26.56K | -- |